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Exoskeleton of an unidentified spider of Borneo

This exoskeleton was found dengeling under a palm leave in one of the river in Borneo on 26-7-2009

 

The exoskeleton of spiders doesn't grow with them, so all spiders have to molt (shed their skin) to grow. They only molt when young, and cease to molt once become an adult.

Spider Molting
Like people, spiders move by contracting muscles attached to a skeleton. But instead of an internal skeleton covered in flesh, spiders have an exoskeleton -- a stiff support structure on the outside of the body. Exoskeleton segments are connected together with joints so the spider can move them back and forth.

Muscles attached on the inside of the exoskeleton contract to move the legs inward, but spiders don't have any muscles to extend the legs back out again. Instead, they have to force bodily fluids (mainly blood) into the legs to push them outward. If a spider loses too much body water, it can't generate the necessary hydraulic pressure to push its legs out. This is why you sometimes see spiders on their backs with their legs curled up.


The exoskeleton is made of several layers of cuticle, a composite material containing various proteins and chitin, a long-chain polysaccharide (sugar). The chitin and protein molecules are arranged in long chains, in successive layers, like the grain in plywood.

This structure makes cuticle extremely strong, as well as highly effective at keeping the spider from drying out, but the material does have one serious drawback. While it's flexible enough for movement, it can't expand like human bones and tissue -- in other words, it can't grow. In order to increase its size, the spider has to form a new, larger cuticle exoskeleton and shed its old one (this is called molting).

Molting occurs frequently when a spider is young, and some spiders may continue to molt throughout their life. At the appropriate time, hormones tell the spider's body to absorb some of the lower cuticle layer in the exoskeleton and begin secreting cuticle material to form the new exoskeleton. The new exoskeleton is typically folded to some extent, so it can expand once the spider sheds the older one. The spider also secretes a molting fluid between the old exoskeleton and the new one. Once the new exoskeleton is finished, the spider absorbs the molting fluid. This creates a gap between the two exoskeletons, which makes it easier to separate them.

Typically, the spider does most of its growing immediately after losing the old exoskeleton, while the new exoskeleton is highly flexible. The new exoskeleton is also very soft in this stage, making the spider particularly vulnerable to attack. Many species will lower themselves on a silk line during the molting process, so they're out of reach of predators while the cuticle material hardens.


Exoskeleton of an unidentified spider of Borneo

Spider Molting

Photo left : exoskeleton of an unidentified spider of Sabah, Borneo Island.

To molt, the spider has to bust out from the inside of this old exoskeleton.

This spider do this by increases heart rate to pump a lot of hemolymph (the spider's blood) from the abdomen into the cephalothorax (the head part). The pressure expands the cephalothorax, which pushes on the old exoskeleton until it cracks. The spider flexes its muscles until this old exoskeleton falls away.

 


View from front
chelicerae (the two jaws)
View from behind
chelicerae (the two jaws)

 

Photo left : Head view of the exoskeleton of the same unidentified spider of Sabah.

From this angle one can see seven (7) of the eight (8) circles of eyes.

Below the eyes is chelicerae (the two jaws).

At the very bottom of the two jaws (out of view from this picture) are  two sharp horizontal fangs.


 

 

 

SPIDERS OF BORNEO

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